Modequai was the father of Morella. The two of them came to Greenwich ten years after the events of The Final Reckoning. Their appearance was quite a shock, as black squirrels were supposedly a dead race. There were rumours that Modequai and his daughter were the last surviving members of the Hawthorn Realm.
Soon after their arrival, Modequai had an audience with the Starwife, Audrey Scuttle. He demanded that the silver acorn be given to his daughter. Though Audrey refused him, Modequai began to gain favour with her grey squirrel subjects, who resented her for being a mouse rather than a squirrel. Eventually, Modequai and a number of grey squirrels barged into the Starwife's chamber and tore the silver acorn from about her neck. Then his daughter became the new Starwife as the 'Usurping Mouse' was chased away.